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Enhanced Facetime: You can now use Facetime over 3G cellular, which means if you have a cellular iPad like me you can now make video calls on the go.
Safari improvements include iCloud Tabs, Safari sync, offline reading, and ability to upload photos and videos to websites from within the browser. This actually is major for me as it looks like I can now post articles to ZDNet all with just the iPad.
Email improvements include better photo and video attachment support, funky new pull to refresh animation, and the ability to assign VIPs to your email. VIPs are synced via iCloud too so they will appear on your iPad and iPhone as well.